has part has part has_part anal vein of the insect wing anterior branch of the beetle wing vein cubital anterior branch 3+4 (CuA3+4) of the beetle wing anal anterior branch 1+2 (AA1+2) of the beetle wing anal anterior branch 1+2 and cubital anterior branch 3+4 (AA1+2+CuA3+4) of the beetle wing The anterior branch of the anal vein that is composed of the anal anterior branch 1+2 and the cubital anterior branch 3+4 of the beetle wing.