develops_from mid culminant late culminant DDANAT:0000011 Dictyostelium_discoideum_anatomy ISBN:0521583640 Kessin book p. 201. Structure in which stalk tube formation has progressed down to the basal disc. The sorogen ascents up the stalk to it apical position during this stage. Terminal differentiation occurs at this stage: death of the stalk cells and encapsulation of the spores. culminant pseudoplasmodium whole organism