has role is protonated form of gallate geroprotector antioxidant trihydroxybenzoic acid gallic acid A trihydroxybenzoic acid in which the hydroxy groups are at positions 3, 4, and 5. 3,4,5-trihydroxybenzoic acid CHEBI:30778 3,4,5-Trihydroxybenzoic acid antineoplastic agent cyclooxygenase 2 inhibitor EC 1.13.11.33 (arachidonate 15-lipoxygenase) inhibitor apoptosis inducer astringent plant metabolite human xenobiotic metabolite