develops from receives synaptic input in region sends synaptic output to region medulla layer M3 medulla layer M4 medulla layer M8 transmedullary Y neuron transmedullary Y neuron TmY4 VFB:FBbt_00003821 TmY4 Pre- versus postsynaptic innervation judged by scoring of terminal morphology from figures in Fischbach and Dittrich (1989) as assessed by FlyBrain Neuron DB. FlyBrain_NDB:10172 Transmedullary Y wide-field neuron that terminates with both fine and bleb-type arborizations in lobula layer 4 and in lobula plate layers 2, 3 and 4, but shows only fine arborizations in lobula plate layer 1. It displays both fine and bleb-type arborizations in medulla layer M8, but has only fine arborizations in medulla layer M3 and M4 (Morante and Desplan, 2008; Fischbach and Dittrich, 1989). It is cholinergic (Konstantinides et al., 2022). fly_anatomy.ontology TmY4 FBbt:00003821 lobula layer 4 lobula plate layer 1 lobula plate layer 2 lobula plate layer 3 lobula plate layer 4 Notch ON hemilineage secondary neuron immature transmedullary Y neuron TmY4 adult cholinergic neuron optic lobe wide-field columnar neuron