lobula plate layer 4 Layer of the lobula plate that is the most distant from the second optic chiasm (Fischbach and Dittrich, 1989). Terminals of T4d and T5d, and most of the vertical system neurons are found in this layer (Fischbach and Dittrich, 1989; Rajashekhar and Shamprasad, 2004). Neurons in this layer are sensitive to downward motion (Henning et al., 2022). layer Lop4 VFB:FBbt_00003889 fly_anatomy.ontology lobula plate layer D FBbt:00003889 FBbt:00040021 lobula plate layer