develops from garland cell garland cell primordium embryonic garland cell fly_anatomy.ontology FBbt:00005704 wreath cell Garland cell of the embryo. This cell is mononucleate, fusing with another embryonic garland cell between stages 13 and 16 to generate the binucleate cells seen in the larva (Zhuang et al., 2009). head mesoderm derivative