develops from meiotic primary spermatocyte primary spermatocyte in meiotic telophase I Make into XP once OBO:rel 'during' is legal. fly_anatomy.ontology Primary spermatocyte during meiotic telophase 1 (GO:0007134). In early telophase the daughter nucleus forms within the karyoplasm and the remnants of karyoplasmic structures. At late telophase, two nuclear membranes have formed around the nucleus of the future secondary spermatocyte, the parafusorial or paranuclear membranes (three to five layers of double membranes) have dissolved and the cell wall has thickened at the site of maximal invagination. FBbt:00057148 primary spermatocyte during late telophase of the first meiotic division primary spermatocyte during early telophase of the first meiotic division primary spermatocyte in meiotic anaphase I