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        <rdf:type rdf:resource="http://www.w3.org/2002/07/owl#TransitiveProperty"/>
        <rdfs:label>part of</rdfs:label>
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    <!-- http://purl.obolibrary.org/obo/BFO_0000051 -->

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        <rdfs:label>overlaps</rdfs:label>
        <rdfs:label xml:lang="en">overlaps</rdfs:label>
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        <rdfs:label>in taxon</rdfs:label>
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    <!-- http://purl.obolibrary.org/obo/NCBITaxon_8459 -->

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        <rdfs:label>Testudines</rdfs:label>
    </Class>
    


    <!-- http://purl.obolibrary.org/obo/UBERON_0000094 -->

    <Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0000094">
        <rdfs:label>membrane organ</rdfs:label>
    </Class>
    


    <!-- http://purl.obolibrary.org/obo/UBERON_0001352 -->

    <Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0001352">
        <rdfs:label>external acoustic meatus</rdfs:label>
    </Class>
    


    <!-- http://purl.obolibrary.org/obo/UBERON_0001691 -->

    <Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0001691">
        <rdfs:label>external ear</rdfs:label>
    </Class>
    


    <!-- http://purl.obolibrary.org/obo/UBERON_0001756 -->

    <Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0001756">
        <rdfs:label>middle ear</rdfs:label>
    </Class>
    


    <!-- http://purl.obolibrary.org/obo/UBERON_0002364 -->

    <Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0002364">
        <rdfs:label>tympanic membrane</rdfs:label>
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        <ns4:IAO_0000115>A thin membrane that separates the external ear from the middle ear, consisting of two epithelia (one part of the the external accoustic meatus epithelium, the other part of the tympanic cavity epithelium) with a fibrous layer between them. Its function is to transmit sound from the air to the ossicles inside the middle ear. The malleus bone bridges the gap between the eardrum and the other ossicles. Rupture or perforation of the eardrum can lead to conductive hearing loss. [WP,unvetted].</ns4:IAO_0000115>
        <oboInOwl:hasDbXref>MESH:D014432</oboInOwl:hasDbXref>
        <oboInOwl:hasDbXref>VHOG:0001146</oboInOwl:hasDbXref>
        <oboInOwl:hasDbXref>XAO:0000212</oboInOwl:hasDbXref>
        <oboInOwl:hasDbXref>FMA:9595</oboInOwl:hasDbXref>
        <oboInOwl:hasDbXref>GAID:870</oboInOwl:hasDbXref>
        <oboInOwl:hasDbXref>UMLS:C0041445</oboInOwl:hasDbXref>
        <oboInOwl:hasExactSynonym>paries membranaceus cavi tympani</oboInOwl:hasExactSynonym>
        <ns4:UBPROP_0000011>The tympanic membrane (TM) has three layers, two epithelia provided by the external acoustic meatus (EAM) and the MEC, derived respectively from the first pharyngeal cleft and pouch, and a mesenchymally derived fibrous stratum</ns4:UBPROP_0000011>
        <oboInOwl:hasDbXref>EMAPA:19062</oboInOwl:hasDbXref>
        <ns4:UBPROP_0000001>The thin membrane which closes externally the cavity of the middle ear and is deeply sited at the bottom of the external auditory meatus. [Dorian_AF, Elsevier&#39;s_encyclopaedic_dictionary_of_medicine, Part_B:_Anatomy_(1988)_Amsterdam_etc.:_Elsevier][VHOG]</ns4:UBPROP_0000001>
        <oboInOwl:id>UBERON:0002364</oboInOwl:id>
        <oboInOwl:hasExactSynonym>membranous wall of tympanic cavity</oboInOwl:hasExactSynonym>
        <oboInOwl:hasDbXref>Wikipedia:Tympanic_membrane</oboInOwl:hasDbXref>
        <oboInOwl:hasDbXref>MA:0000257</oboInOwl:hasDbXref>
        <ns4:UBPROP_0000003>In particular, a tympanic membrane was not present in the earliest tetrapods (...). An ear utilizing a tympanic membrane evolved independently at least three times in tetrapods: (1) in the lineage that leads to anurans (frogs), (2) in the line of evolution to turtles and diapsids, and (3) in the late synapsid lineage that gave rise to mammals.[well established][VHOG]</ns4:UBPROP_0000003>
        <oboInOwl:hasExactSynonym>ear drum</oboInOwl:hasExactSynonym>
        <oboInOwl:hasExactSynonym>eardrum</oboInOwl:hasExactSynonym>
        <oboInOwl:hasDbXref>NCIT:C12502</oboInOwl:hasDbXref>
        <oboInOwl:hasExactSynonym>myrinx</oboInOwl:hasExactSynonym>
        <oboInOwl:hasExactSynonym>Rivinus&#39; membrane</oboInOwl:hasExactSynonym>
        <ns4:UBPROP_0000012>most AOs consider this part of the middle ear, so we go with this and place it in the middle ear as a border to the external, although it could be considered to overlap the external ear in that it has an EAM layer as part - for this reason we do not consider middle and external ear to be spatially disjoint.</ns4:UBPROP_0000012>
        <oboInOwl:hasDbXref>SCTID:181180005</oboInOwl:hasDbXref>
        <oboInOwl:hasExactSynonym>lateral wall of tympanic cavity</oboInOwl:hasExactSynonym>
        <oboInOwl:hasRelatedSynonym>tympanum</oboInOwl:hasRelatedSynonym>
        <ns4:RO_0002161 rdf:resource="http://purl.obolibrary.org/obo/NCBITaxon_8459"/>
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    </Class>
    


    <!-- http://purl.obolibrary.org/obo/UBERON_0004114 -->

    <Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0004114">
        <rdfs:label>tympanic cavity</rdfs:label>
    </Class>
    


    <!-- http://purl.obolibrary.org/obo/UBERON_0005042 -->

    <Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0005042">
        <rdfs:label>inner epithelial layer of tympanic membrane</rdfs:label>
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    <!-- http://purl.obolibrary.org/obo/UBERON_0009213 -->

    <Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0009213">
        <rdfs:label>pharyngeal membrane of 1st arch</rdfs:label>
    </Class>
    


    <!-- http://purl.obolibrary.org/obo/UBERON_0010069 -->

    <Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0010069">
        <rdfs:label>outer epithelial layer of tympanic membrane</rdfs:label>
    </Class>
    


    <!-- http://purl.obolibrary.org/obo/UBERON_0010070 -->

    <Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0010070">
        <rdfs:label>intermediate layer of tympanic membrane</rdfs:label>
    </Class>
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<!-- Generated by the OWL API (version 3.2.4.1806) http://owlapi.sourceforge.net -->



