abnormal sporulation abnormal nuclear separation after meiosis abnormal separation of nuclei in ascus abnormal nuclear separation in ascus abnormal nuclear separation after meiotic nuclear division FYPO:0002711 A cellular process phenotype in which the haploid nuclei formed by meiotic nuclear division do not separate normally. Nuclei may be closer together than normal, or remain attached by an incompletely separated outer nuclear membrane, or both. abnormal nuclear distribution after meiosis 2013-09-25T14:53:56Z abnormal nuclear separation during sporulation midori