glucuronosyltransferase activity N-acetylglucosaminyl-proteoglycan 4-beta-glucuronosyltransferase activity https://github.com/geneontology/go-ontology/issues/28762 https://github.com/geneontology/go-ontology/issues/30193 GO:0050509 Catalysis of the reaction: 3-O-{alpha-D-GlcNAc-[(1->4)-beta-D-GlcA-(1->4)-alpha-D-GlcNAc](n)-(1->4)-beta-D-GlcA-(1->3)-beta-D-Gal-(1->3)-beta-D-Gal-(1->4)-beta-D-Xyl}-L-seryl-[protein] + UDP-alpha-D-glucuronate = 3-O-{[(1->4)-beta-D-GlcA-(1->4)-alpha-D-GlcNAc](n+1)-(1->4)-beta-D-GlcA-(1->3)-beta-D-Gal-(1->3)-beta-D-Gal-(1->4)-beta-D-Xyl}-L-seryl-[protein] + H+ + UDP. Reactome:R-HSA-9036285 UDP-alpha-D-glucuronate:N-acetyl-alpha-D-glucosaminyl-(1->4)-beta-D-glucuronosyl-proteoglycan 4-beta-glucuronosyltransferase activity N-acetylglucosaminylproteoglycan beta-1,4-glucuronyltransferase activity RHEA:20908 Reactome:R-HSA-3656267 N-acetylglucosaminyl-proteoglycan 4-b-glucuronosyltransferase activity molecular_function Reactome:R-HSA-2022856 heparan glucuronyltransferase II activity EC:2.4.1.225 Reactome:R-HSA-9036289 MetaCyc:2.4.1.225-RXN Reactome:R-HSA-2076392 Reactome:R-HSA-3656257