cytoplasmic actin-based contraction involved in cell motility cytoplasmic actin-based contraction involved in forward cell motility The actin filament-based movement by which cytoplasmic actin filaments slide past one another resulting in a contraction that propels the cell in the direction that has been defined as the front of the cell. biological_process cytoplasmic actin-based contraction involved in forward cell locomotion GO:0060328