oxidoreductase activity, acting on paired donors, with oxidation of a pair of donors resulting in the reduction of molecular oxygen to two molecules of water palmitoyl-[glycerolipid] 3-(E)-desaturase activity https://github.com/geneontology/go-ontology/issues/28070 https://github.com/geneontology/go-ontology/issues/28362 MetaCyc:RXN-8319 MetaCyc:RXN-8318 MetaCyc:RXN-1725 RHEA:46764 molecular_function 1-18:1-2-16:0-phosphatidylglycerol trans-3 desaturase activity MetaCyc:RXN-16053 EC:1.14.19.43 GO:0102654 Catalysis of the reaction: a 1-acyl-2-hexadecanoyl-glycerolipid + 2 H+ + O2 + 2 reduced [2Fe-2S]-[ferredoxin] = a 1-acyl-2-[(3E)-hexadec-3-enoyl]-glycerolipid + 2 H2O + 2 oxidized [2Fe-2S]-[ferredoxin]. The this activity introduces an unusual trans double bond at carbon 3 of a palmitoyl group attached to the sn-2 position of glycerolipids.