p-2S-SMAD2/3:SMAD4:PARP1 p-2S-SMAD2/3:SMAD4 PARP1 binds SMAD2/3:SMAD4 heterotrimer Authored: Orlic-Milacic, M, 2012-04-04 Edited: Jassal, B, 2012-04-10 PARP1 (poly [ADP-ribose] polymerase 1) binds SMAD2/3:SMAD4 heterotrimers associated with DNA SMAD-binding elements (SBEs). Pubmed21095583 Reactome Database ID Release 432187330 Reactome, http://www.reactome.org ReactomeREACT_121212 Reviewed: Huang, Tao, 2012-05-14