Ub-SMAD4 Ub-SMAD4:USP9X USP9X (FAM) binds to ubiquitinated SMAD4 Authored: Williams, MG, 2010-06-08 Edited: Jassal, B, 2012-04-10 In the cytosol, a ubiquitin hydrolase USP9X (FAM) binds to ubiquitinated SMAD4 (Dupont et al. 2009). Pubmed19135894 Reactome Database ID Release 43870479 Reactome, http://www.reactome.org ReactomeREACT_121227 Reviewed: Huang, Tao, 2012-05-14