has part has part transferred as integrative and conjugative element ICE conjugation module T4SS-type integrative and conjugative element Meng LIU (Mia), Oliver He http://db-mml.sjtu.edu.cn/ICEberg/ T4SS-type ICE a typical class of ICEs which transfer as single stranded DNA molecules and require T4SS and can both found in Gram-positive bacteria and Gram-negative bacteria ICE integration and excision module single stranded deoxyribonucleic acid