has_part has part has_part has_part Brucella lipopolysaccharide Brucella O-polysaccharide rough Brucella strain YL the rough Brucella strain is Brucella that has a rough lipopolysaccharide as its part. (rough lipopolysaccharide is lack of surface expression or reduced surface expression of the O polysaccharide ) rough Brucella lipopolysaccharide Brucella