part_of adult columella adult columella stalk Fig. 12-4, in Biology of Ticks ISBN:0-19-505910-7. TADS:0000349 TADS:0000349 The thin section of cuticle that connects the columella to the underside of the cuticular layer that supports the pedicels. The columella extends from the upper lip of the macula to the underside of the pedicels. default_namespace