develops from forerunner cell group A group of cells that migrate at the leading edge of shield during gastrulation but do not involute. At the end of gastrulation, forerunner cells migrate deep into the embryo and organize to form Kuppfer's vesicle. Cooper and D'Amico, 1996. DFCs TAO:0000023 dorsal forerunner cells forerunner cells teleost_anatomy noninvoluting endocytic marginal cell cluster embryonic structure