develops in develops in epithelium scleral mesenchyme epithelial scleral papilla layer Scleral papillae (which induce scleral ossicles in birds) have not been reported in teleosts, but evidence for their presence may not have been sought UBERON:0010300 scleral papillae The scleral ossicles are of dermal (neural crest) origin, develop intramembranously within the scleral mesenchyme, and are induced to form by overlying (transient) epithelial scleral papillae (Nelson, 1942; Murray, 1943; Fyfe and Hall, 1981, 1983; Hall, 1981, 2005; Pinto and Hall, 1991)