cerebellar granule layer morphology trait cerebellar glomerulus morphology trait Any measurable or observable characteristic related to the shape, structure, color, or pattern of the intertwined cluster of nerve fibers surrounded by glia where mossy fibers from the pontine nuclei in the white matter synapse with granule cell axons, Golgi cell axons and unipolar brush interneuron axons. Trait.ontology VT:0004099